“…With the amelioration of climate about 9,000 years ago, forest began to expand and riparian zones were gradually established [Gunin et al 1999]. This environment did not support mammoths, rhinoceros, ostriches, or even, apparently, the archaic horse [Orlova et al 2004], and thus they disappeared; but it did support aurochs and the gradual appearance of elk, ibex, bear, and, in some riparian zones, moose, and boar [Jacobson-Tepfer 2013]. At the end of the middle Holocene and with the retreat of forest cover, horses (takhi) had reappeared on the steppe; but by three thousand years ago even aurochs and moose disappeared and other species began to pull back to the higher elevation.…”