“…The nearly complete separation of the Mediterranean from the Atlantic Ocean during the late Miocene (Martín et al, 2001(Martín et al, , 2009(Martín et al, , 2010Betzler et al, 2006;Flecker et al, 2015;Tulbure et al, 2017;Capella et al, 2018;Ng et al, 2021) had a global impact on the thermohaline ocean circulation and, consequently, on the climate and organic matter fluxes (Pérez-Asensio et al, 2012, 2014Jiménez-Moreno et al, 2013). In the Betic Cordillera, several straits connected the Mediterranean and the Atlantic throughout the Guadalquivir Basin (Martín et al, 2001(Martín et al, , 2009(Martín et al, , 2014Betzler et al, 2006;Puga-Bernabéu et al, 2022).…”