“…Besides these groups, the Coleoptera fauna was hardly studied until the 2000s. More intensive research has been carried out using a variety of methods in the reserve over the past 12 years (Kurmaeva et al 2008, Egorov et al 2010, Feoktistov 2011, Egorov and Ruchin 2012, 2013a, 2013b, 2014, Pavlov and Ruchin 2013, Legalov et al 2014, Egorov et al 2015, Egorov 2017, Sazhnev 2017, Ruchin and Egorov 2018a, 2018d, 2018e, 2019b, Tomaszewska et al 2018, Kazantsev et al 2019, 2019c. There are published data on individual families: Carabidae (Feoktistov 2008, Staphylinidae (Semenov 2014, Kurbatov and Egorov 2014, Semenov 2015, Scydmaenidae (Kurbatov and Egorov 2014), Curculionidae (Scolytinae) (Mandelshtam and Egorov 2017.…”