“…Although the grazer-alga food chain is important, it is not the sole structuring force in tropical rocky shores; it can be affected by other factors such as oceanographic forcing like ENSO (El Niño Southern Oscillation) cycles (Vinueza et al 2006), by seasonal cycles (Williams 1993a(Williams , 1993b(Williams , 1994, number of recruits (Kaehler and Williams 1997), predators (Menge et al 1986a) and complex interactions (Sauer Machado et al 1996). In temperate rocky shores, grazers are also affected by nutrient subsidies (Bustamante et al 1995), the availability of recruits (Freidenburg et al 2007), desiccation and high winds (Bazterrica et al 2007), disturbance such as storms, sand and ice scour (McQuaid and Dower 1990;Chapman 1991, 1993;Underwood 1999), upwelling and oceanographic conditions (Menge et al 2003).…”