“…In addition, browse under forest crowns is less likely to be buried by snow (Harestad, 1985), and some conifers such as Douglas-fir (Pseudotsuga menziesii) and western redcedar (Thuja plicata) are fed on by ungulates as are the arboreal lichens attached to their boles and canopy (Dawson et al, 1990;Pauley et al, 1993;Armleder et al, 1994;D'Eon, 2001). Overall, however, there is far less browse under canopies than open habitats (Jameson, 1967;Thomas, 1979;Peek et al, 2001;Sullivan et al, 2007). Ungulates, therefore, must weight the varied benefits of a forest canopy against the costs of an overall lower abundance of browse under dense canopies (Jameson, 1967;Thomas, 1979;Peek et al, 2001;Sullivan et al, 2006Sullivan et al, , 2007.…”