The crustal structure of north-central Wyoming records a history of complex lithospheric evolution from Precambrian accretion to Cretaceous-Paleogene Laramide shortening. We present two active source P wave velocity model profiles collected as part of the Bighorn Arch Seismic Experiment in 2010. Analyses of these velocity models and single-fold reflection data, together with potential field modeling of regional gravity and magnetic signals, constrain crustal structure and thickness of the Bighorn region. We image a west dipping reflection boundary and model a sharp magnetic contact east of the Bighorn Arch that together may delineate a previously undetected Precambrian suture zone. Localized patches of a high-velocity, high-density lower crustal layer (the "7.× layer") occur across the study area but are largely absent beneath the Bighorn Arch culmination. Moho topography is relatively smooth with no large-scale offsets, with depths ranging from~50 to 37 km, and is largely decoupled from Laramide basement topography. These observations suggest that (1) the edge of the Archean Wyoming craton lies just east of the Bighorn Mountains, approximately 300 km west of previous interpretations, and (2) Laramide deformation localized in an area with thin or absent 7.× layer, due to its relatively weak lower crust, leading to detachment faulting. Our findings show that Precambrian tectonics in northern Wyoming may be more complicated than previously determined and subsequent Laramide deformation may have been critically dependent on laterally heterogeneous crustal structure that can be linked to Precambrian origins.The potential role of Archean (>2.5 Ga) and Proterozoic (0.541-2.5 Ga) structures in the development of the Laramide orogeny (~75 Ma to~40 Ma) has been suggested by numerous studies that note contrasting structural patterns in the southern and northern Rocky Mountain provinces [Brown, 1993;Erslev and Koenig, 2009;Stone, 1986]. Deformation, probably driven by flat-slab subduction [e.g., Bird, 1988;Coney and Reynolds, 1977;Dickinson and Snyder, 1978], led to basement-involved thick-skinned shortening during the Laramide orogeny (~75 Ma to~40 Ma) [Dickinson et al., 1988], forming basement-cored foreland arches. In the Proterozoic basement of Colorado, Laramide arches typically strike north-northwest, are separated by narrow basins, and form a well-defined, en echelon deformation front on their eastern margin. In the large Archean basement of Wyoming, Laramide structures exhibit a larger variety of strikes and are separated by wide basins, with no well-defined deformation front. These contrasting geometries of thick-skinned deformation have been attributed to the greater diversity of structural fabrics in the Archean Wyoming basement [Brown, 1993;Erslev and Koenig, 2009] and different crustal architectures. Most notably, the high-velocity, high-density lower crustal WORTHINGTON ET AL.CRUSTAL STRUCTURE OF THE BIGHORN ARCH 208