Research on the dispositional origins of political preferences is flourishing, and the primary conclusion drawn from this work is that stronger needs for security and certainty attract people to a broad-based politically conservative ideology. Though this literature covers much ground, most integrative assessments of it have paid insufficient attention to the presence and implications of contingencies in the relationship between dispositional attributes and political attitudes. In this article, we review research showing that relationships between needs for security and certainty and political preferences vary considerably-sometimes to the point of directional shifts-on the basis of (1) issue domain and (2) contextual factors governing the content and volume of political discourse individuals are exposed to. On the basis of this evidence, we argue that relationships between dispositional attributes and political preferences vary in the extent to which they reflect an organic functional resonance between dispositions and preferences or identity-expressive motivation to adopt a political attitude merely because it is discursively packaged with other need-congruent attitudes. We contend that such a distinction is critical to gaining a realistic understanding of the origins and nature of ideological belief systems, and we consequently recommend an increased focus on issue-based and contextual variation in relationships between dispositions and political preferences.KEY WORDS: political preferences, ideology, needs for security and certainty, political expertise Why do individuals' political preferences lean to the left or to the right? To put it mildly, a great amount of social-scientific ink has been spilled addressing this question. Some perspectives suggest that political preferences are a function of interests associated with the positions of the social groups one belongs to, with members of more powerful groups adopting more conservative views (e.g., Bobo, 1999;Huber & Form, 1973;Kluegel & Smith, 1986; Marx & Engels, 1846/1970Runciman, 1966;Sears & Funk, 1991;Sidanius & Pratto, 1999;Weeden & Kurzban, 2014). Other approaches have focused more closely on social relationships, with a sizable body of research suggesting that people adopt the political attitudes and beliefs that are normative in the reference groups with which they identify (Berelson, Lazarsfeld, & McPhee, 1954;Merton, 1957;Newcomb, 1943) or common within their families or social networks (Huckfeldt, Johnson, & Sprague, 2004;Jennings & Niemi, 1981; Advances in Political Psychology, Vol. 39, Suppl. 1, 2018 doi: 10.1111 bs_bs_banner Jost, Ledgerwood, & Hardin, 2008;Sears & Levy, 2003). Last but not least, research in political science zeroes in on the role of political parties, arguing that individuals acquire the preferences enunciated by the leaders of the parties they identify with (Campbell, Converse, Miller, & Stokes, 1960;Goren, 2005;Lenz, 2013;Zaller, 1992). Approaches like these share a common thread: They focus on the social or contextua...