This paper reports on the design, modeling, analysis, and evaluation of a micro-electromechanical systems acoustic sensor and the novel design of an acoustic vector sensor array (AVS) which utilized this acoustic sensor. This research builds upon previous work conducted to develop a small, lightweight, portable system for the detection and location of quiet or distant acoustic sources of interest. This study also reports on the underwater operation of this sensor and AVS. Studies were conducted in the lab and in the field utilizing multiple acoustic sources (e.g., generated tones, gun shots, drones). The sensor operates at resonance, providing for high acoustic sensitivity and a high signal-to-noise ratio (SNR). The sensor demonstrated a maximum SNR of 88 dB with an associated sensitivity of −84.6 dB re 1 V/μPa (59 V/Pa). The sensor design can be adjusted to set a specified resonant frequency to align with a known acoustic signature of interest. The AVS demonstrated an unambiguous, 360-degree, in-plane, azimuthal coverage and was able to provide an acoustic direction of arrival to an average error of within 3.5° during field experiments. The results of this research demonstrate the potential usefulness of this sensor and AVS design for specific applications.