“…However, seedlings are often less tolerant of herbivory than mature plants (Strauss and Agrawal, 1999;Colpetzer et al, 2004). Also, the effect of herbivore damage can be enhanced by the addition of other environmental stresses (Berg et al, 2015;Harris, 1981). For example, defoliation can have a more severe effect on shaded plants relative to unshaded plants or on species that may be less shade tolerant (Baraza et al, 2004;Blundell and Peart, 2001; Lentz and Cipollini, 1998;Norghauer et al, 2008;Rogers and Siemann, 2003).…”