“…To date, 63 quantitative trait loci (QTLs) for FLA have been identified based on linkage analysis, including 9, 7, 6, 4, 4, 5, 4, 7, 6, 0, 8, and 3 on chromosomes 1–12, respectively ( Supplementary Table 1 ) (Li et al, 1999 ; Yan et al, 1999 ; Dong et al, 2003 ; Kobayashi et al, 2003 ; Luo et al, 2008 ; Zhang et al, 2008 , 2013 ; Cai, 2009 ; Hu et al, 2012 ; Wang et al, 2012 ; Bian et al, 2014 ; Zou et al, 2014 ; Zhu et al, 2016 ; Ham et al, 2019 ). In addition, 100 QTLs for FLA have been detected based on association analysis (Huang et al, 2010 ; Chen et al, 2012 ; Lu et al, 2015 ; Dong H. J. et al, 2018 ; Dong Z. Y. et al, 2018 ). Among these QTLs, only one QTL qFla-8-2 has been fine-mapped and predicted the candidate genes (Zhu et al, 2016 ).…”