“…However, they must have co-existed alongside earlier Homo species for over 1 million years (Oms et al, 2000;Toyo-Moyano et al, 2013), with macaques and hominins found at a number of Early and Middle Pleistocene sites including Cueva Negra del Estrecho del Quípar and Solana del Zamborino in Spain, and Galeria Pesada in Portugal (see Marigó et al, in press). Evidence of hominins (both fossils of H. neanderthalensis and/or stone tools) is also present with Macaca (although not always within the same levels) at a number of Late Pleistocene sites, including Hunas (Rosendahl et al, 2011), Grotta degli Orsi Volanti (Mazza et al, 2005), Kugelsteinhöhle II (Fladerer, 1991), Cova Negra (Marigó et al, in press) and Lezetxiki (Castanos et al, 2011). Modern ecological studies indicate that Barbary macaques do not live comfortably alongside humans, and unlike baboons, vervets and some Asian macaques, do not exploit human habitats (Menard, 2002).…”