http://www.eje.cz tained only from nestedness will not provide clear explanations if environmental factors are not associated with this process (Murkami & Hirao, 2010). Thus, using environmental variables to account for the pattern of nestedness recorded for island ecosystems will enable conservationists to focus on the factors associated with diversity and protect the islands harbouring most species.Diff erent processes can generate patterns in nestedness: (1) passive sampling: can generate nestedness because common species are more accessible and thus easy to sample than rare species (Andrén, 1994) and (2) selective extinction of species: species inhabiting ecosystems experiencing species loss, would disappear from sites in a predictable way, which could lead to nestedness (Cutler, 1991; Lomolino, 1996), (3) diff erential colonization of habitats: the nested structure of isolated habitats could be due to dispersal limitations as species diff er in their ability to colonize (Ryti & Gilpin, 1987; Cook, 1995; Lomolino, 1996), (4) habitat nestedness: considers the nestedness of species assemblages to be a consequence of their close association with habitats that have a nested distribution (Ward & Lakhani, 1977; Honnay et al., 1999).