“…The relatively simple horn cores with relatively faint keel-like ridges appear plesiomorphic, more like living boselaphins and unlike most Old World fossil tragoportacins (Bibi, 2009, Bibi et al, 2009) from the later Miocene such as Miotragocerus , Austroportax , Sivaceros , Sivoreas , Protragocerus , Kipsigicerus , and Tragoportax that normally have long, strongly compressed horn cores characterized by anterior step-like indentations and well-formed keels (Solounias and Moelleken, 1992). Even some species of Miotragocerus , a genus with diverse morphology that probably requires revision (Hartung et al, 2020:281), that have relatively simple, compressed shorter horns, such as M. monacensis , M. pannoniae , or M. abyssinicus , exhibit greater differences in degree of horn core compression, demarcations, prominent anterior keels, posterior groove(s), no posterior curvature, and no raised ridge between horn cores (see Spassov and Geraards, 2004; Bibi, 2011; Fuss et al, 2015; Hartung et al, 2020). Other Miocene “boselaphins” with cores exhibiting distinct keels, strong mediolateral compression, great divergence, and/or torsion differ from Neotragocerus and include Pachyportax , Plioportax , Samokeros , Selenoportax , and Sivaportax .…”