Abstract:Authors' Contribution NM conceived and designed the study. NM and TM analysed the data and wrote the manuscript. PR improved the manuscript through valuable discussion and constructive comments. AA and SM participated in the fieldwork.
“…The semi‐selenodont pattern of the molar confirms the inclusion of this specimen in family Tragulidae (Colbert, 1935; Draz, Ni, et al, 2020; Farooq, Khan, & Akhtar, 2007; Morales, Soria, Sánchez, Quiralte, & Pickford, 2003; Pickford, 2001). Dorcatherium was represented by four Siwalik species D. minimus , D. nagrii , D. minus , and Dorcatherium majus (Colbert, 1935; Farooq, Khan, Akhtar, & Khan, 2007).…”
Section: Systematic Palaeontologysupporting
confidence: 62%
“…One is the large‐sized G. superba and the other is small‐sized Gazella lydekkeri (Pilgrim, 1937, 1939). The presence of strong ribs, strong stylids, and the absence of median basal pillar in upper molars, are the principal morphometric characters of G. lydekkeri (Draz, Ni, et al, 2020; Draz, Samiullah, et al, 2020; Samiullah, Naseem, et al, 2015). The morphological difference of characters between the two species is: the teeth in G. superba are larger as compared to G. lydekkeri ; the presence of rudimentary entostyle in G. superba (Saeed, Khan, Iqbal, & Amin, 2018; Samiullah, Naseem, et al, 2015) and relative prominence of anterior and posterior median ribs in G. superba (Khan et al, 2013b; Khan et al, 2013a).…”
Section: Systematic Palaeontologymentioning
confidence: 99%
“… The studied specimens (Referred data is taken from Akhtar, 1992; Ali, Khan, & Akhtar, 2012; Abbas et al, 2018; Aftab, Ahmad, Khan, & Akhtar, 2015; Aftab et al, 2019; Bhatti, Khan, & Akhtar, 2012; Bhatti, Khan, Akhtar, Khan, et al, 2012; Batool, Khan, & Qureshi, 2015; Draz, Ni, et al, 2020; Draz, Samiullah, et al, 2020; Farooq, Khan, Akhtar, & Khan, 2008; Iqbal, Khan, Atiq, Ikram, & Akhtar, 2011; Khan et al, 2010, 2013; Khan et al, 2015; Saeed et al, 2018;Samiullah et al, 2011; Samiullah, Naseem, et al, 2015; Samiullah, Jabeen, et al, 2015; Zaheer, Mahmood, Babar, & Akbar, 2017). …”
Section: Systematic Palaeontologymentioning
confidence: 99%
“…A detailed description of the geological and biostratigraphical settings of the Siwaliks was provided by Barry et al (2002). The large mammalian faunas from the Siwalik Group has been the focus of many researchers due to their biostratigraphic and paleoenvironmental implications (Abbas et al, 2018; Akhtar, Samiullah, Rasul, & Altaf, 2020; Bernor, Feibel, & Viranta, 2003; Draz, Ni, et al, 2020; Draz, Samiullah, et al, 2020; Grégoire et al, 2009; Grégoire, Chaimanee, Jaeger, & Ducrocq, 2001; Grégoire, Guo, & Beard, 2004; Grégoire, Moine, O'reilly, Cottin, & Giret, 2000; Samiullah, Akhtar, Khan, & Ghaffar, 2012; Samiullah, Naseem, et al, 2015; Samiullah, Jabeen, et al, 2015; Scott, 1940).…”
Section: Introductionmentioning
confidence: 99%
“…Location map of Potwar Plateau in Northern Pakistan: showing the studied locality Dhok Bun Ameer Khatoon (enboxed), Chinji Formation (highlighted) in district Chakwal, Northern Pakistan with generalized stratigraphic section of the major Siwalik Group formations (Draz, Ni, et al, 2020; Draz, Samiullah, et al, 2020; Khan et al, 2017) [Colour figure can be viewed at wileyonlinelibrary.com]…”
The fossil site Dhok Bun Ameer Khatoon (DBAK) is located 35 km southeast from Chakwal district, Punjab, Pakistan. It has yielded a significant amount of fossil artiodactyls including five families: Bovidae, Tragulidae, Suidae, Giraffidae, and Anthracotheriidae. This site principally exposes Middle Miocene strata that have been dated approximately as 14.2–11.63 Mya. A detailed study was carried out on the fossils found at DBAK and 29 specimens have been described herein. The collection comprises isolated upper and lower teeth and fragments of mandibular ramii. Gazella lydekkeri, Elachistoceras khauristanensis, Dorcatherium majus, Listriodon pentapotamiae, Giraffokeryx punjabiensis, Giraffa priscilla, and Merycopotamus nanus have been described in this article. The newly discovered material is productive to understand the fauna of this Middle Miocene locality and it enhances our knowledge about the described species. Merycopotamus nanus have been reported for the first time from this locality. Palaeoenvironmental data indicates that the Miocene climate of Pakistan was most likely to be monsoonal. All the feeding habits and distribution of those animals suggested the presence of abundant forests, herbaceous, grassy, and bushy vegetation in the DBAK and its surroundings.
“…The semi‐selenodont pattern of the molar confirms the inclusion of this specimen in family Tragulidae (Colbert, 1935; Draz, Ni, et al, 2020; Farooq, Khan, & Akhtar, 2007; Morales, Soria, Sánchez, Quiralte, & Pickford, 2003; Pickford, 2001). Dorcatherium was represented by four Siwalik species D. minimus , D. nagrii , D. minus , and Dorcatherium majus (Colbert, 1935; Farooq, Khan, Akhtar, & Khan, 2007).…”
Section: Systematic Palaeontologysupporting
confidence: 62%
“…One is the large‐sized G. superba and the other is small‐sized Gazella lydekkeri (Pilgrim, 1937, 1939). The presence of strong ribs, strong stylids, and the absence of median basal pillar in upper molars, are the principal morphometric characters of G. lydekkeri (Draz, Ni, et al, 2020; Draz, Samiullah, et al, 2020; Samiullah, Naseem, et al, 2015). The morphological difference of characters between the two species is: the teeth in G. superba are larger as compared to G. lydekkeri ; the presence of rudimentary entostyle in G. superba (Saeed, Khan, Iqbal, & Amin, 2018; Samiullah, Naseem, et al, 2015) and relative prominence of anterior and posterior median ribs in G. superba (Khan et al, 2013b; Khan et al, 2013a).…”
Section: Systematic Palaeontologymentioning
confidence: 99%
“… The studied specimens (Referred data is taken from Akhtar, 1992; Ali, Khan, & Akhtar, 2012; Abbas et al, 2018; Aftab, Ahmad, Khan, & Akhtar, 2015; Aftab et al, 2019; Bhatti, Khan, & Akhtar, 2012; Bhatti, Khan, Akhtar, Khan, et al, 2012; Batool, Khan, & Qureshi, 2015; Draz, Ni, et al, 2020; Draz, Samiullah, et al, 2020; Farooq, Khan, Akhtar, & Khan, 2008; Iqbal, Khan, Atiq, Ikram, & Akhtar, 2011; Khan et al, 2010, 2013; Khan et al, 2015; Saeed et al, 2018;Samiullah et al, 2011; Samiullah, Naseem, et al, 2015; Samiullah, Jabeen, et al, 2015; Zaheer, Mahmood, Babar, & Akbar, 2017). …”
Section: Systematic Palaeontologymentioning
confidence: 99%
“…A detailed description of the geological and biostratigraphical settings of the Siwaliks was provided by Barry et al (2002). The large mammalian faunas from the Siwalik Group has been the focus of many researchers due to their biostratigraphic and paleoenvironmental implications (Abbas et al, 2018; Akhtar, Samiullah, Rasul, & Altaf, 2020; Bernor, Feibel, & Viranta, 2003; Draz, Ni, et al, 2020; Draz, Samiullah, et al, 2020; Grégoire et al, 2009; Grégoire, Chaimanee, Jaeger, & Ducrocq, 2001; Grégoire, Guo, & Beard, 2004; Grégoire, Moine, O'reilly, Cottin, & Giret, 2000; Samiullah, Akhtar, Khan, & Ghaffar, 2012; Samiullah, Naseem, et al, 2015; Samiullah, Jabeen, et al, 2015; Scott, 1940).…”
Section: Introductionmentioning
confidence: 99%
“…Location map of Potwar Plateau in Northern Pakistan: showing the studied locality Dhok Bun Ameer Khatoon (enboxed), Chinji Formation (highlighted) in district Chakwal, Northern Pakistan with generalized stratigraphic section of the major Siwalik Group formations (Draz, Ni, et al, 2020; Draz, Samiullah, et al, 2020; Khan et al, 2017) [Colour figure can be viewed at wileyonlinelibrary.com]…”
The fossil site Dhok Bun Ameer Khatoon (DBAK) is located 35 km southeast from Chakwal district, Punjab, Pakistan. It has yielded a significant amount of fossil artiodactyls including five families: Bovidae, Tragulidae, Suidae, Giraffidae, and Anthracotheriidae. This site principally exposes Middle Miocene strata that have been dated approximately as 14.2–11.63 Mya. A detailed study was carried out on the fossils found at DBAK and 29 specimens have been described herein. The collection comprises isolated upper and lower teeth and fragments of mandibular ramii. Gazella lydekkeri, Elachistoceras khauristanensis, Dorcatherium majus, Listriodon pentapotamiae, Giraffokeryx punjabiensis, Giraffa priscilla, and Merycopotamus nanus have been described in this article. The newly discovered material is productive to understand the fauna of this Middle Miocene locality and it enhances our knowledge about the described species. Merycopotamus nanus have been reported for the first time from this locality. Palaeoenvironmental data indicates that the Miocene climate of Pakistan was most likely to be monsoonal. All the feeding habits and distribution of those animals suggested the presence of abundant forests, herbaceous, grassy, and bushy vegetation in the DBAK and its surroundings.
This article provides a detailed taxonomic study of mammalian fossil fauna from five localities situated within the Middle Siwalik subgroup including the Nagri and Dhok Pathan formations in Punjab, Pakistan. Twenty‐three euungulate specimens comprised of isolated teeth, and maxillary and mandibular fragments, are described. This collection includes the bovid, Elachistoceras; a very rare faunal element in the Siwaliks of Pakistan, as well as Elachistoceras khauristanensis, Pachyportax latidens, Giraffa punjabiensis, Bramatherium grande, Merycopotamus dissimilis, Dorcatherium minus, Dorcatherium majus, Hippopotamodon sivalense, Sivalhippus theobaldi, Sivalhippus nagriensis and Brachypotherium perimense These fossil remains add important new insights into the taxonomy and diversity of Late Miocene mammal faunas of the Middle Siwaliks. The data is important for understanding the biogeographical and palaeoenvironmental history of the region. The characteristics of the fossils described in this study further support the currently hypothesised presence of a massive open land environment with variable wet and dry seasons alike to that of the current climate in Eurasia and Africa. The variable habitat niches of these co‐existing fauna also give further support to the supposition that there was a much more mixed array of palaeoenvironments ranging from a prevalence of woodland to expansive savannah territory during the deposition of Nagri and Dhok Pathan formations.
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