“…Hence, the occurrence of C. zekuensis from Zeku indicates that this area had a more moderate paleoclimate with higher temperature (especially coldest month mean temperature (CMMT)) and precipitation in the Miocene than today, and the co‐occurring broadleaved woody plants (e.g., megathermic Cercis and Koelreuteria Laxmann, mesothermic Acer L., Fraxinus L., and Ulmus L.; Guo, 1980; Li et al, 2016) corroborate the existence of a warm temperate forest during the early Miocene. Rather higher historical biodiversity reflected by fossil insects (e.g., Cixius Latreille, Wesmaelius Krüger, and Helius Lepeletier & Serville; Li et al, 2017b; Yang et al, 2018; Wu et al, 2019), spiders, and birds (e.g., Passer Brisson; Young, 1975) also support this deduction. Furthermore, the inferred paleoecological trait appears to match very well the coeval regional or global climate changes from other independent proxies (e.g., geobiomarkers or isotope records; Zachos et al, 2001; Liu et al, 2014).…”