New Species of Tipulidae From the Paleocene of Central Alberta, Canada

Abstract

Three new species of fossil Tipulidae are described. Pseudolimnophila speirsae n. sp. and Limnophila marklae n. sp. (subfamily Limoniinae, tribe Hexatomini) are represented by adults; Tipula tubifera n. sp. (subfamily Tipulinae, tribe Tipulini) is represented by a pupa. Six other pupae belong to the subfamily Tipulinae.

The Gomphaeschninae (Odonata: Aeshnidae): new fossil genus, reconstructed phylogeny, and geographical history

“…The fossil aeshnids found to date include six exuviae of immature stages, not described here, and are associated with many other adult and larval insects (Wighton, 1982), aquatic and terrestrial plants, and at Site 4 bones of freshwater fishes. A warm-temperate to subtropical climate and low-energy , shallow, fresh water associated with mixed forest is envisaged (Wighton, 1980(Wighton, , 1982. Modern gomphaeschnines are strong fliers, breeding in lakes and streams and living near and flying over water, mainly in warm climates (Lieftinck, 1968;Westfall, 1978); their life-history agrees with other information available about Palaeocene habitats in Alberta.…”

“…The Palaeocene fossils are unusual when compared with other Early Tertiary insect faunas of North America (Wilson, 1978(Wilson, , 1982 because they have a relatively low diversity of Diptera and Hymenoptera, and include both adult and immature stages of a variety of higher taxa (Mitchell & Wighton, 1979;Wighton, 1982). Of the taxa represented, Tipulidae (Wighton, 1980) and Orthoptera (Kevan & Wighton, 1981) have been described in detail.…”

New fossil Tipulidae from the volcano-sedimentary Latest Oligocene of Bes-Konak (Turkey)

“…The numerous finds of fossil representatives of this group come from Europe, from where more than 60 species have been described by HEER (1849), NOVAK (1877), MEUNIER (1899MEUNIER ( , 1906MEUNIER ( , 1915MEUNIER ( , 1916, LOEW (1850), COCKERELL (1917COCKERELL ( , 1921, COCKERELL & HAINES (1921), HENRIKSEN (1922), ALEXANDER (1931), STATZ (1934STATZ ( , 1944, THEOBALD (1937), ZEUNER (1938), FREIWALD (1990), GENTILINI (1990) and KRZEMIŃSKI (2000), and from North America by SCUDDER (1890SCUDDER ( ,1894, HANDLIRSCH (1910), ALEXANDER (1938), MELANDER (1949), LEWIS (1969LEWIS ( , 1973 and WIGHTON (1980), who described over 30 species, mainly from the genus Tipula LINNAEUS, 1758.…”

A new species of Tipula LINNAEUS, 1758 (Diptera: Tipulidae) from Green River Formation, USA

“…Currently the family Tipulidae comprises over 4000 extinct and extant species (EVENHUIS 1994(EVENHUIS , 1996OOSTERBROEK 2018), but the knowledge about the origin and early evolution of this group of insects is still relatively scarce. (Table 1; SCUDDER 1877SCUDDER , 1890SCUDDER , 1894COCKERELL 1909COCKERELL , 1910HANDLIRSCH 1910;ALEXANDER 1938;MELANDER 1949;LEWIS 1973;WIGHTON 1980;BROWN 1985;EVENHUIS 1994). The largest number of fossil species of Tipula were described from Florissant, Colorado (Eocene-Oligocene).…”

“…Four species were described from the Oligocene (Ruby River Basin in Montana, Fossil Canyon in Utah, and White River in Colorado). Only two species of Tipula were described from the Eocene: WIGHTON, 1980) was described from the Paleocene of Alberta (Table 1). From the Paleogene, over 70 species are known, Palearctic representatives of Tipula were described from Denmark (HENRIKSEN 1922;FREIWALD 1990), Turkey (KANIA & NEL 2013), and Baltic Region (LOEW 1850;MEUNIER 1899MEUNIER , 1906ALEXANDER 1931).…”