“…Their shoot development pattern is categorized as a flush and succeeding-type leaf emergence (Kikuzawa, 1983); they flush shoots generally once in spring, sometimes flushing new shoots if environmental conditions are preferable, such as gap formation. According to Yamaguchi et al (2011), both the oak species are relatively O 3 tolerant among the Japanese tree species and are related to a noted capacity for isoprene emission (Loreto and Fares, 2007;Tani and Kawawata, 2008;Miyama et al, 2013). Although there have been several studies primarily investigating the effects of elevated CO 2 and/or O 3 on leaf physiological and morphological traits in oak species (e.g., Velikova et al, 2005;Paoletti et al, 2007;Watanabe et al, 2007Watanabe et al, , 2013, only a few studies have investigated the combined effects of elevated CO 2 and O 3 on the growth and carbon allocation in oak species (Quercus petraea, Broadmeadow and Jackson, 2000;cf.…”