“…Aerodynamic effects that predominate on insect wings, such as the lift produced by wing rotation and vortex shedding, are intrinsically included in this model, as these mechanisms produce aerodynamic force and are not intrinsically different from other sources of lift and drag (Walker, 2002). For instance, periodic vortex shedding produces tonal sound, but not as a separate acoustic mechanism, because the way vortex shedding produces sound is by producing periodic cycles of lift and drag on the solid object (Blake, 2017a;Blake, 2017b, chapter 4). This mechanism provides a general explanation for why the flight tone in certain directions is, in many species, loudest (dominant) at the fundamental frequency of the wingbeat, while in other directions 2014), who used wing kinematics of a male Ruby-throated hummingbird (Archilochus colubris, body mass: 2.7 g, wingbeat frequency: 57 Hz) and a computational fluid dynamics (CFD) model to calculate normalized force components for a single flapping hummingbird wing, in the i=x, y and z directions.…”