“…They are considered ecosystem engineers, altering soil composition and hydrology (Jones et al, 1994;Mando et al, 1996) and are the dominant invertebrate decomposer in these environments (Collins, 1981;Holt, 1987;Schuurman, 2005), playing an important role in nutrient cycling (Holt and Coventry, 1990;Lepage et al, 1993;Konaté et al, 1999). They are also sensitive to moisture and temperature levels (AbenspergTraun and De Boer, 1990;Smith and Rust, 1994;Korb and Linsenmair, 1998), and thus expected to respond to seasonal variation.…”