“…N saturation (defined as N availability in excess of biological demand) has been cited as a contributor to forest mortality (Aber et al, 1989;Magill et al, 2000Magill et al, , 2004. High levels of forest floor N, relative to biological demand, can lead to foliar nutrient imbalance (Zoettl and Huettl, 1986;Cronan and Grigal, 1995), reduced red spruce cold tolerance (Sheppard, 1994;Schaberg et al, 2002), and increased freeze injury to foliage that in turn reduce carbohydrate reserves and increase the potential for secondary stressors that can cause tree mortality (DeHayes, 1992;Schaberg et al, 1997;Perkins et al, 2000). Other environmental stressors, such as drought (Johnson and Siccama, 1983;Driscoll et al, 2003) and aluminum toxicity in roots (Shortle and Smith, 1988), or nutrient leaching from the forest floor (Berg, 1986;Johnson et al, 1994;Adams, 1999) could act synergistically with N saturation to increase the potential for tree mortality on already pollutant-stressed forests.…”