[1] The vertical structure and evolution of the wintertime annular modes are studied using 6 years of geopotential height, carbon monoxide (CO), and water vapor (H 2 O) data from Aura Microwave Limb Sounder. The Northern Hemisphere annular mode (NAM) and the Southern Hemisphere annular mode (SAM) reveal a strong coupling of the dynamics in the stratosphere and mesosphere between 316 hPa (∼9 km) and 0.002 hPa (∼90 km). CO is a good tracer throughout the middle atmosphere, while variable vertical gradients of H 2 O limit the regions where it is useful as a dynamical tracer. The maximum of the CO NAM and SAM (CNAM and CSAM) indices is used to monitor and characterize the evolution of wintertime polar dynamics as a function of time and height. The CNAM analysis reveals reformation of a stronger mesospheric polar vortex after significant stratospheric sudden warmings in 2006, 2009, and 2010. There is a significant anticorrelation between the mesospheric and stratospheric CNAM indices during 2005-2010 winters, supporting the hypothesis of mesosphere-stratosphere coupling through planetary-gravity wave interactions.