“…In this paper we only consider incompressible velocity fields: ∇ • v = 0, where v = (v x , v y ), and ∇ = (∂/∂x, ∂/∂y). Indeed, two-dimensional velocity fields in the oceans are incompressible to a very good approximation, although in some local regions vertical currents can alter this picture [25], potentially impacting population dynamics [13,26,27].…”