“…As the largest family, and most well studied HSP, the HSP20 family of proteins is ATP-independent and generally assemble into large oligomers that can protect other proteins from denaturation and aggregation ( Papsdorf & Richter, 2014 ; Asea, Kaur & Calderwood, 2016 ). An increasing body of evidence has shown that HSP20s regulate the responses of plants to environmental changes and thus allow plants to survive adverse conditions ( Sun et al, 2012 , 2016 , 2020 ; Murakami et al, 2004 ; Charng et al, 2006 ; Shakeel, Heckathorn & Luthe, 2012 ; Khurana, Chauhan & Khurana, 2013 ; Zou et al, 2012 ; Kaur et al, 2015 ; Chauhan et al, 2012 ; Wang et al, 2020 ; Mu et al, 2013 ; Li et al, 2018 ). In a previous study, we identified HSP20 genes within the genome of watermelon and found that HSP20s in plants can be divided into 18 subfamilies ( He et al, 2019 ).…”