“…They have proved useful in several areas of the biological sciences. Perhaps the most obvious application is in population genetics; implicitly in the early work of Fisher and Wright [18], and explicitly in later developments [19,20,21,22,23,24,25]. However they have also been used in a number of other areas such as the study of radioactive particles in animals [26,27,28], the study of patterns of vegetation [29], models of interaction between species [30,31,32] and metapopulation models [33].…”