“…The formation is divided into three members: the Lower Member is composed of thin bioclastic limestones, muddy limestones and shales; the Middle Member is mainly made up of fine pyroclastic rocks with a few sandy and muddy limestones; and the Upper Member consists of calcareous clastic rocks with a small amount of bioclastic limestones ( Hou et al, 1993 ). The formation is mostly Famennian in age ( Ma et al, 2017 ; Zong et al, 2020 ; Shen et al, 2021 ; Stachacz et al, 2021 ). The Hongguleleng Formation is very rich in many types of the early Famennian fossils, such as acritarchs, bivalves, brachiopods, bryozoans, cephalopods, chondrichthyans, conodonts, conulariids, corals, echinoderms, gastropods, ostracods, plants, radiolarians, spores, trace fossils, and trilobites ( Liao, 2002 ; Zong R W, 2021, unpublished specimens).…”