The La Chilca Formation is exposed at the Poblete Norte and Ancha sections, Talacasto, Central Precordillera. Detailed facies analysis on field outcrops suggests a tide-dominated tidal plain model on the Silurian siliciclastic ramp, integrated by supratidal to subtidal plain deposits. The lower part of the La Chilca Formation (upper supratidal plain) is herein attributed to reworked glaciomarine deposits that occurred during interglacial Hirnantian age, under conditions of sea rise and the action of tidal amplification. The intertidal-supratidal depositional environments are represented by tidal channels, scour and fill, tidal dunes, and hummocky cross-stratified sandstones. The estimated tidal conditions for the tidal plain were mesotidal to macrotidal, under action of combined flows, attributed to the interaction of tidal currents and orbital low-velocity waves. The identification of microbially induced sedimentary structures (MISS) in these deposits, such as petee polygonal structures, pitted texture, imploded sand volcanoes, Astropolithon sp., Manchuriophycus sp., domal build-ups, biofilms, elephant skin, and trace fossils such as Chondrites sp., Zoophycos sp., Undichnia sp., Planolites sp., Monomorphichnus sp., and gastropod traces, may have played a protection and stabilization role on the clastic sediment surfaces against erosion of the La Chilca Formation deposits. The prolific growth of microbial mat on the intertidal to supratidal zone occurs explicitly at the turning points of transgression-regression cycles in shallow-water environments under stable base-level conditions. The middle-upper section of the La Chilca Formation shows non-cyclic rhythmites that are replaced by the cyclic rhythmites, suggesting an open tidal plain subject to an orbital/climatic rather than a tectonic control of cyclicity (Milankovitch cycles). The aggradation in this section indicates a shallowing towards the top of the formation. The La Chilca Formation records two third-order transgressive events and a shallowing event. The transgressive events are found in the base and top of the formation and are marked by the presence of Fe-oolites, while the shallowing event is determined by the presence of MISS.