“…The number of these species is less than those found by Siwi (2017) on the Beach Si Runtoh, Baluran National Park, East Java (19 species); Supusepa (2018) in Negeri Tial and Negeri Suli, Maluku (30 and 39 species, respectively); or even Rumahlatu and Leiwakabessy (2017) in the coastal waters of Ambon Island, Maluku (65 species). However, the number of species in the forest is still greater than that reported by Ernanto et al (2010) in Muara Sungai Batang, South Sumatra (10 species), Isnaningsih and Patria (2018) in Tanjung Lesung, Banten (7 species), Maturbongs and Elviana (2016) in Kambapi Coast, Papua (10 species), Nurrudin et al (2015) in Tanjung Jabung Barat, Jambi (15 species), Rangan (2010) in Minahasa, North Sulawesi (9 species), and Puryono and Suryanti (2019) in Pemalang, Central Java (8 species). Species density at LT-1 is higher than LT-2 density and so subsequently density at LT-2 compared to LT-3.…”