IntroductionStudies of seasonal changes of the nitrogenous and carbohydrate fractions of many plants have been made, but in few cases have the plants been sufficiently divided to show the functions of individual tissues. Usually when detailed work has been done, not all parts have been analyzed. Of course, many plants are constructed so as to make subdivision difficult and impractical. Hence, in this study it was decided to use the common cultivated sunflower (Helianthus annutus L.) as it fulfilled the requirement of being easily divided into anatomically distinct regions.Some work has already been done on various tissues of this plant over short periods. CLEMENTS (12) has dealt with the hourly variations of carbohydrates in the leaves. He has likewise studied seasonal changes in the sunflower, and although these data have not been published, they will be referred to from time to time in this paper.The purpose of this work was to determine the functions of plant tissues during various stages of growth and to obtain information of the synthesis and translocation of organic solutes.