“…Although, this temperate-tropical example of bryophyte flora (Dierßen 2001) is distributed across Europe, tropical Africa, and southern, southeastern and southwestern parts of Asia, its populations are rather scattered (Porley 2013, Hodgetts 2015). The European scope of the range includes: to the north Denmark, to the west Great Britain, in the south France and Italy, in central areas Austria, Belgium, the Czech Republic, Germany, Hungary, the Netherlands, Poland, Slovakia and Switzerland, in the southeast Bulgaria, Romania, Slovenia, Serbia and Turkey, and in the east Belarus, the Crimea, Latvia, northwest Russia and Ukraine (Papp et al 2013, ECCB 2016. In the majority of these countries it has a defined status category, and it was recently included in the European Red List of Mosses, Liverworts and Hornworts (Hodgetts et al 2019).…”