“…During land plant evolution, the expansion of SNARE proteins and their functional diversification was hypothesized to be linked to multicellularity and probably facilitated adaptation to a terrestrial lifestyle (Sanderfoot, 2007). Thus, land plant secretory pathways are highly sophisticated, dynamic and diversely regulated, being involved in a manifold of cellular processes ranging from polarized growth to defence responses (Batoko & Moore, 2001; Collins et al ., 2003; Catalano et al ., 2007; Enami et al ., 2009; Silva et al ., 2010; Reichardt et al ., 2011; Uemura et al ., 2012; Ichikawa et al ., 2014; Johansson et al ., 2014; Yun et al ., 2016; Xia et al ., 2019; Hirano et al ., 2020; Rubiato et al ., 2021). In M. polymorpha , the SYP1 protein family consists of four members: SYP12A, SYP12B, SYP13A and SYP13B.…”