“…During the late Pleistocene, the North Pacific developed into a cold, rocky shore ecosystem that provided complex and productive environments for many marine species, and dynamic ocean currents and upwelling that led to higher plankton productivity (Paine 2002, Graham et al 2003, Lyle et al 2010. Coastal areas in the North Pacific are highly variable environments, where changes in current patterns, ocean circulation, and upwelling have reversed in different time scales, ranging from 10,000 to 100,000 yr BP (Kennett and Ingram 1995, Kiefer et al 2001, Hewitt 2004, affecting productivity (Webb and Bartlein 1992).…”