“…The ebb and flow of the tides would be expected to resuspend some pollen, and potentially cause sorting according to pollen size and buoyancy (Brush and Brush, 1972;Holmes, 1990Holmes, , 1994. Several studies have suggested that pollen grains in suspension in water behave as fine silt-size particles (Brush and Brush, 1972;Clark, 1986;Farr, 1989;Holmes, 1994), although some contribute together with mineral matter to larger particles as the result of flocculation/agglomeration, to be expected in a saline environment, or are incorporated into faecal pellets (Pickrill, 1987;Chmura and Eisma, 1995;Chmura et al, 1999).…”