Pragmatic Classification of the World's Salticidae (Araneae)

Abstract

This paper, dedicated to search for identification methods of genera of Salticidae (Araneae), presents prototype of a “Handbook of Jumping Spiders Identification”, based on morphology of palps, spermathecae and ducts, as well as some other easily noticeable characters. It includes diagnostic drawings of representative species of each genus, additional survey of diversity of these characters in 4800 recognizable species is available instantly, by hyperlinks provided to parallel Internet "Monograph of the Salticidae (Araneae) of the World 1995-2016".Part I "Introduction to alternative classification of Salticidae" by Prószyński (2016a), accessible at: http://www.peckhamia.com/salticidae/Subfamilies/ [too large to be published whole as a PDF]. The work contains methodological suggestions on how the proposed system could be improved and further developed. Partial revision of the present taxonomic system of Salticidae is included.The paper provides diagnoses and diagnostic drawings to genera of Salticidae, grouped to facilitate identification into morphologically coherent, informal groups of genera. There are following provisional groups proposed: AEURILLINES, AMYCINES, AMYCOIDA VARIA, ASTIAINES, BELIPPINES, CHRYSILLINES, COCALODINES, COLONINES [= former Thiodininae], DENDRYPHANTINES, DIOLENINES, EUODENINES, EUOPHRYINES, EUPOAINES, EVARCHINES, HABRONATTINES, HARMOCHIRINES, HELIOPHANINES, HISPONINES, HYLLINES, ICIINES, LAPSIINES, LIGONIPEINES, LYSSOMANINES, MENEMERINES, MYRMARACHNINES, NOTICIINES, PELLENINES, PSEUDICIINES, SIMAETHINES, SITTICINES, SPARTAEINES, THIRATOSCIRTINAE, YAGINUMAELLINES, YLLENINES. There is also temporary UNCLASSIFIED group and display of exemplary FOSSILS. The proposals of grouping and delimitation have working character, pending further research and tests.The following synonyms and combinations (new, corrected or reinstated) are listed in the paper together with their documentation and/or discussions. They have been accumulated during 22 years of work on database, but are printed for the first time only now (location of their documentation in the text below can be quickly found using computer searching facility).Aelurillus stanislawi (Prószyński, 1999) (male from Israel) = Rafalus stanislawi Prószyński, 1999, Aelurillus stanislawi Azarkina, (2006) (nec Prószyński, 1999) = Aelurillus minutus Azarkina, 2002, Amphidraus manni (Bryant 1943) = Nebridia manni Bryant 1943, Amphidraus mendica (Bryant 1943) = Nebridia mendica Bryant 1943, Amphidraus semicanus (Simon, 1902) = Nebridia semicana Simon, 1902, Bianor incitatus Thorell, 1890 (in part) = Stichius albomaculatus Thorell, 1890, Bryantella smaragdus (Crane, 1945) = Bryantella smaragda (Crane, 1945), Chinattus undulatus (Song & Chai, 1992) (in part, male) = Chinattus szechwanensis (Prószyński, 1992), Colyttus kerinci (Prószyński & Deeleman-Reinhold, 2012) = Donoessus kerinci Prószyński & Deeleman-Reinhold, 2012, Colyttus nigriceps (Simon, 1899) = Donoessus nigriceps (Simon, 1899), Colyttus striatus (Simon, 1902) = Donoessus striatus (Simon, 1902), Cytaea severa (Thorell, 1881) (in part) = Cytaea alburna Keyserling, 1882, Euophrys minuta Prószynski, 1992 ) = Lechia minuta (Prószynski, 1992 ), Laufeia daiqini (Prószyński & Deeleman-Reinhold, 2012) = Junxattus daiqini Prószyński & Deeleman-Reinhold, 2012, Laufeia kuloni (Prószynski & Deeleman-Reinhold, 2012) = Orcevia kuloni Prószynski & Deeleman-Reinhold 2012, Laufeia keyserlingi (Thorell, 1890) = Orcevia keyserlingi (Thorell, 1890), Laufeia eucola (Thorell, 1890) = Orcevia eucola (Thorell, 1890), Laufeia perakensis (Simon, 1901) = Orcevia perakensis (Simon, 1901), Laufeia proszynskii Song, Gu & Chen, 1988 = Orcevia proszynskii (Song, Gu & Chen, 1988), Laufeia squamata ( Żabka, 1985 ) = Lechia squamata Żabka, 1985, Maevia C. L. Koch, 1846 (in part) = Paramaevia Barnes, 1955, Maevia hobbsae Barnes, 1958 = Paramaevia hobbsae Barnes, 1958, Maevia michelsoni Barnes, 1958 = Paramaevia michelsoni (Barnes, 1958), Maevia poultoni Peckham & Peckham, 1909 = Paramaevia poultoni (Peckham & Peckham, 1901),Maratus anomaliformis (Żabka, 1987) = "Lycidas" anomaliformis Żabka, 1987, Metaphidippus felix (Peckham & Peckham, 1901) = Messua felix (Peckham & Peckham, 1901), Monomotapa principalis Wesolowska, 2000 = Iranattus principalis (Wesolowska, 2000), Myrmarachne exasperans (Peckham & Peckham, 1892) = Emertonius exasperans Peckham & Peckham, 1892, Myrmarachne melanocephala MacLeay, 1839 (in part) = Myrmarachne ramosa Badcock, 1918, Myrmarachne melanocephala MacLeay, 1839 (in part) = Myrmarachne contracta (Karsch, 1880), Myrmarachne melanocephala MacLeay, 1839 (in part) = Myrmarachne albicrurata Badcock, 1918, Myrmarachne melanocephala MacLeay, 1839 (in part) = Myrmarachne lateralis Badcock, 1918, Myrmarachne melanocephala MacLeay, 1839 (in part) = Myrmarachne providens Simon, 1901, Myrmavola globosa (Wanless, 1978) = Toxeus globosus (Wanless, 1978) (self-correction), Omoedus albertisi (Thorell, 1881) = Zenodorus albertisi (Thorell, 1881), Omoedus arcipluvii (Peckham, Peckham, 1901) = Zenodorus arcipluvii (Peckham, Peckham, 1901), Omoedus asper (Karsch, 1878) = Ascyltus asper (Karsch, 1878), Omoedus bernsteini (Thorell, 1881) = Zenodorus bernsteini (Thorell, 1881), - Omoedus brevis Zhang J., Maddison, 2012 = Zenodorus brevis (Zhang J., Maddison, 2012), Omoedus cyanothorax (Thorell, 1881) = Pystira cyanothorax (Thorell, 1881), - Omoedus durvillei (Walckenaer, 1837) = Zenodorus durvillei (Walckenaer, 1837)- Omoedus danae (Hogg, 1915) = Zenodorus danae Hogg, 1915, - Omoedus darleyorum Zhang J., Maddison, 2012 = Zenodorus darleyorum (Zhang J., Maddison, 2012),Omoedus ephippigerus (Simon, 1885) = Pystira ephippigera (Simon, 1885), Omoedus karschi (Thorell, 1881) = Pystira karschi (Thorell, 1881), Omoedus lepidus (Guerin, 1834) = Zenodorus lepidus (Guerin, 1834), Omoedus metallescens (Koch L., 1879) = Zenodorus metallescens (Koch L., 1879), Omoedus meyeri Zhang J., Maddison, 2012 = Zenodorus meyeri (Zhang J., Maddison, 2012), Omoedus microphthalmus (Koch L., 1881) = Zenodorus microphthalmus (Koch L., 1881), Omoedus nigripalpis (Thorell, 1877) = Pystira nigripalpis (Thorell, 1877)]. Omoedus obscurofemoratus (Keyserling, 1883) = Zenodorus obscurofemoratus (Keyserling, 1883), Omoedus omundseni Zhang J., Maddison, 2012 = Zenodorus omundseni (Zhang J., Maddison, 2012), Omoedus orbiculatus (Keyserling, 1881) = Zenodorus orbiculatus (Keyserling, 1881), Omoedus papuanus Zhang J., Maddison, 2012 = Zenodorus papuanus (Zhang J., Maddison, 2012), Omoedus ponapensis (Berry, Beatty, Prószyński, 1996) = Zenodorus ponapensis Berry, Beatty, Prószynski, 1996, Omoedus semirasus (Keyserling, 1882) = Zenodorus semirasus (Keyserling, 1882), Omoedus swiftorum Zhang J., Maddison, 2012 = Zenodorus swiftorum (Zhang J., Maddison, 2012), Omoedus tortuosus Zhang J., Maddison, 2012 = Zenodorus tortuosus (Zhang J., Maddison, 2012), Omoedus versicolor (Dyal, 1935) = Pystira versicolor Dyal, 1935, [Unrecognizable species of Zenodorus: Omoedus jucundus (Rainbow, 1912) = Zenodorus jucundus (Rainbow, 1912), Omoedus juliae (Thorell, 1881) = Zenodorus juliae (Thorell, 1881), Omoedus marginatus (Simon, 1902) = Zenodorus marginatus (Simon, 1902), Omoedus niger (Karsch, 1878) = Zenodorus niger (Karsch, 1878), - Omoedus pupulus (Thorell, 1881) = Zenodorus pupulus (Thorell, 1881), - Omoedus pusillus (Strand, 1913) = Zenodorus pusillus (Strand, 1913), Omoedus rhodopae (Hogg, 1915) = Zenodorus rhodopae (Hogg, 1915), Omoedus syrinx (Hogg, 1915) = Zenodorus syrinx Hogg, 1915, Omoedus variatus (Pocock, 1899) = Zenodorus variatus (Pocock, 1899), Omoedus varicans (Thorell, 1881) = Zenodorus varicans Thorell, 1881, Omoedus wangillus (Strand, 1911) = Zenodorus wangillus Strand, 1911], Pellenes ostrinus (Simon, 1884) (in part) = Pellenes diagonalis Simon, 1868, Pseudicius alter Wesolowska, 1999 = Afraflacilla altera (Wesolowska, 1999), Pseudicius arabicus (Wesolowska, van Harten, 1994) = Afraflacilla arabica Wesolowska, van Harten, 1994, Pseudicius bipunctatus Peckham, Peckham, 1903 = Afraflacilla bipunctata (Peckham, Peckham, 1903), Pseudicius braunsi Peckham, Peckham, 1903 = Afraflacilla braunsi (Peckham, Peckham, 1903), Pseudicius datuntatus Logunov, Zamanpoore, 2005= Afraflacilla datuntata (Logunov, Zamanpoore, 2005), Pseudicius elegans (Wesolowska, Cumming, 2008) = Afraflacilla elegans (Wesolowska, Cumming, 2008), Pseudicius eximius Wesolowska, Russel-Smith, 2000 = Afraflacilla eximia (Wesolowska, Russel-Smith, 2000), Pseudicius fayda Wesolowska, van Harten, 2010 = Afraflacilla fayda (Wesolowska, van Harten, 2010), Pseudicius flavipes Caporiacco, 1935 = Afraflacilla flavipes (Caporiacco, 1935), Pseudicius histrionicus Simon, 1902 = Afraflacilla histrionica (Simon, 1902), Pseudicius imitator Wesolowska, Haddad, 2013 = Afraflacilla imitator (Wesolowska, Haddad, 2013), Pseudicius javanicus Prószynski, Deeleman-Reinhold, 2012 = Afraflacilla javanica (Prószynski, Deeleman-Reinhold, 2012), Pseudicius karinae (Haddad, Wesolowska, 2011) = Afraflacilla karinae (Haddad, Wesolowska, 2011), Pseudicius kraussi Marples, 1964 = Afraflacilla kraussi (Marples, 1964), Pseudicius mikhailovi Prószynski, 1999 = Afraflacilla mikhailovi (Prószynski, 1999), Pseudicius mushrif Wesolowska, van Harten, 2010 = Afraflacilla mushrif (Wesolowska, van Harten, 2010), Pseudicius philippinensis Prószynski, 1992 = Afraflacilla philippinensis (Prószynski, 1992), Pseudicius punctatus Marples, 1957 = Afraflacilla punctata (Marples, 1957), Pseudicius refulgens Wesolowska, Cumming, 2008 = Afraflacilla refulgens (Wesolowska, Cumming, 2008), Pseudicius reiskindi Prószynski, 1992 = Afraflacilla reiskindi (Prószynski, 1992), Pseudicius roberti Wesolowska, 2011 = Afraflacilla roberti (Wesolowska, 2011), Pseudicius spiniger (Pickard-Cambridge O., 1872) = Afraflacilla spiniger (Pickard-Cambridge O., 1872), Pseudicius tamaricis Simon, 1885 = Afraflacilla tamaricis (Simon, 1885), Pseudicius tripunctatus Prószynski, 1989 = Afraflacilla tripunctata (Prószynski, 1989), Pseudicius venustulus Wesolowska, Haddad, 2009 = Afraflacilla venustula (Wesolowska, Haddad, 2009), Pseudicius wadis Prószynski, 1989 = Afraflacilla wadis (Prószynski, 1989), Pseudicius zuluensis Haddad, Wesolowska, 2013 = Afraflacilla zuluensis (Haddad, Wesolowska, 2013), Servaea incana (Karsch, 1878) (in part) = Servaea vestita ( L. Koch, 1879), Sidusa extensa (Peckham & Peckham, 1896) = Cobanus extensus (Peckham & Peckham, 1896), Sidusa Peckham & Peckham, 1895 (in part) = Cobanus F. O. Pickard-Cambridge , 1900, Sidusa Peckham & Peckham, 1895 (in part) = Wallaba Mello-Leitão, 1940, Stagetillus elegans (Reimoser, 1927) = "Padillothorax" elegans Reimoser, 1927, Stagetillus taprobanicus (Simon, 1902) = "Padillothorax" taprobanicus Simon, 1902, Telamonia besanconi (Berland & Millot, 1941) = Brancus besanconi (Berland & Millot, 1941), Telamonia fuscimana (Simon, 1903) = Brancus fuscimanus (Simon, 1903), Telamonia longiuscula (Thorell, 1899) = Hyllus longiusculus (Thorell, 1899), Telamonia thoracica (Thorell, 1899) [="Viciria"thoracica: Prószyński, 1984 = Hyllus thoracicus (Thorell, 1899), - Thiania sundevalli (Thorell, 1890) = Nicylla sundevalli Thorell, 1890, Thiania spectrum (Simon, 1903) = Thianitara spectrum Simon, 1903, Thiania thailandica (Prószyński & Deeleman-Reinhold, 2012) = Thianitara thailandica Prószyński & Deeleman-Reinhold, 2012, Viciria albocincta Thorell, 1899 = Hyllus albocinctus (Thorell, 1899), Yaginumaella striatipes (Grube, 1861) (in part) = Yaginumaella ususudi Yaginuma, 1972.

Sitticine jumping spiders: phylogeny, classification, and chromosomes (Araneae, Salticidae, Sitticini)

“…Prószyński's more recent (2017a) classification into genera, however, is discordant in many respects with our results, as can be seen in the many combinations that we establish or reinstate below in order to achieve monophyly of genera and subgenera. This discord may have arisen partly because Prószyński was not attempting to create a taxonomy that reflected phylogenetic relationships, but rather the distribution of a few diagnostic characters (Prószyński 2017b).…”

“…Prószyński, who developed our understanding of north-temperate species in a series of papers (1968,1971,1973,1980), recently (2016, 2017a) partitioned this diversity into several genera: Sittipub Prószyński, 2016, Sittiflor Prószyński, 2017, Sittilong Prószyński, 2017, Sittisax Prószyński, 2017, Sittiab Prószyński, 2017, Attulus, and Sitticus. Prószyński did not intend this classification to be phylogenetic, but rather "pragmatic" (Prószyński 2017b), which is to say, not based on a conceptual justification. If a classification rejects reference to a broader theory, whether about monophyly or adaptive zones or predictivity across many characters, then it is not clear what it means, how it can be tested, or whether it can even be correct, except in its specific statements about the few characters mentioned.…”

Delimitation of Mediterranean Genus Iberattus Gen. N., With Comments on Genus Saitis (Araneae: Salticidae)

“…The paper follows methodology of "pragmatic classification" proposed by Prószyński (2016aPrószyński ( , 2017 and practiced by Prószyński, Lissner, & Schäfer (2018) "Taxonomic survey of the genera Euophrys, Pseudeuophrys and Talavera, with description of Euochin gen. n. (Araneae: Salticidae) and with proposals of a new research protocol", Ecologica Montenegrina 18: 24-74. which postulates identification of Salticidae by direct comparison of either specimens, drawings and photographs (whichever is possible), complemented now by experience of Jørgen Lissner, Aart Noorden, Pierre Oger and by Michael Schäfer in macrophotography of live spiders, which revolutionizes diagnostics but requires deep change in collecting procedures. The present paper is based on data of literature, supported by own experience of authors, with a shift to color macrophotographs of live specimens, whenever possible.…”

“…The paper analyses two little know European species of EUOPHRYINES, listed now by World Spider Catalog as Euophrys semiglabrata (Simon, 1868) and Saitis imitatus (Simon, 1868). Their current classification is based on general appearance and shape of palps, the latter being characteristic for the whole group of genera EUOPHRYINES (according to Prószyński (2017), (Euophryini according to Maddison, (2015)), but devoid of characters linking them with known genera. Current genus placement cannot be sustained, however, because both species differ from the above mentioned genera by the most important genus character -internal structure of epigyne (Figs 1Q,N respectively).…”

Review of Genera Evarcha and Nigorella, With Comments on Emertonius, Padilothorax, Stagetillus, and Description of Five New Genera and Two New Species (Araneae: Salticidae)

“…Stagetillus semiostrinus Prószyński, 1987: 104-105 (mf). Stagetillus semiostrinus Prószyński, 2017b: 129, f . 56L (f).…”