“…This local action of trees and beavers varied over the recent centuries, during which humans removed trees and heavily hunted beavers until protection and restoration resulted in recovery from these human actions (e.g., Montgomery, 1997;Butler and Malanson, 2005). Beyond this complication by variation over time, many interrelated variables affect the geomorphic action of riparian trees (e.g., sediment supply and size, nature of bank material, type and age of riparian vegetation, particularly of the root system; Hähne, 1982;Pflug, 1982;Montgomery, 1997) and beavers (e.g., water depth, nature of bank material, tree canopy cover; Broschart et al, 1989;Gurnell, 1998). Further complications relate to interactions between the riparian vegetation and beavers.…”