“…In aquatic environment, mannans in algal fronds and their detritus are consumed not only by microorganisms but also by benthic animals such as annelids, arthropods and mollusks. These invertebrates also possess mannandegrading enzymes and assimilate the mannans as a carbon and energy sources (Chuang and Yang, 1991;Xu et al, 2002a;Song et al, 2008;Ootsuka et al, 2006;Zahura et al, 2010). For complete depolymerization of mannan, at least two types of mannan-degrading enzymes are necessary.…”