“…The Lashkerak and Ghelli Formations provide an integrated ichnologic, sedimentologic, and geobiologic record of the Ordovician Radiation, which represents a changeover point in the history of marine ecosystems (Mángano et al, 2016; Mángano & Droser, 2004; Sepkoski, 1995; Servais & Harper, 2018; Servais et al, 2010; Stigall et al, 2019). The increase in ichnodiversity and degree and depth of bioturbation that took place in marine environments during the Ordovician Radiation had major geobiologic impacts, including most notably the further disruption of microbially stabilized surfaces (Bayet‐Goll, Knaust, et al, 2021; Bayet‐Goll, Uchman, et al, 2021; Buatois & Mángano, 2012; Buatois et al, 2016; Buatois et al, 2020; Mángano et al, 2016; Mángano & Droser, 2004). In particular, bioturbation by large, relatively deep, highly mobile deposit feeders, including the so‐called sediment bulldozers, was particularly effective in sediment mixing, being detrimental to the formation of matgrounds.…”