“…Improve nutrient use efficiency Nutrient cycling Soil N and P explained 13.5% and 12.4% of crop yield variances, respectively, at the Karst CZO (Liang et al, 2020); high risk of environmental pollution prevents increasing crop yield through fertilization (Liang et al, 2021) -Shift from "leaky" to "tight" N-cycling processes (D. Li et al, 2018) -Re-emergence of microbial N-cycling functions in soil after agricultural abandonment (D. Li et al, 2018Li et al, , 2021a 15 Farmers in all CZs identified fertilisers as their greatest cost and one of the top three farming challenges they face (Zheng et al,. 2018, this paper) 1,2 -Reduce N inputs to improve economic livelihoods and well-being -Judicious use of N with crop residue returns increases soil C (X. D. Song et al, 2019a;X. W. Song et al, 2019b) -Reduce fertilizer use to increase crop yields (Liang et al, 2020(Liang et al, , 2021 Reduce point-source pollution by reducing fertiliser use…”