experiment with pigeons, each trial began with a fixedinterval (FI) 5-sec schedule on a white center key (the search period). When a pigeon completed the FI schedule, the choice period began. The center key remained white, and a side key was lit either green or red (with equal probability); these two key colors represented two different types of prey that were associated with two different handling times. The green key represented a prey item with a short handling time (an FI 5-sec schedule leading to food), and the red key represented a prey item with a long handling time (an FI 20-sec schedule leading to food). With either alternative, the pigeon could accept the prey (by pecking the green or red key) or reject it (by pecking the white key three times, which would return the pigeon to the search period and the beginning of a new FI 5-sec schedule). The pigeons almost always accepted the short alternative (the green key), which is not surprising, because rejecting it and returning to the search period would always increase the delay to the next food delivery. Of greater theoretical interest were the pigeons' choices when the long alternative (the red key) was presented.To predict whether the pigeons would accept or reject the long alternative, Lea (1979) relied on a version of optimal foraging theory now known as optimal diet theory (Cook & Cockrell, 1978;Sih & Christensen, 2001). Stated simply, optimal diet theory predicts that the animal should accept the long alternative only if, on average, it provides a greater amount of food per unit time than could be obtained by returning to the search period (cf. Goss-Custard, 1977;Krebs, Erichsen, Webber, & Charnov, 1977;Snyderman, 1983). For this specific example, optimal diet theory states that the pigeon should always reject the long alternative, because a strategy of continually returning to the search period until the short alternative (the green key) is presented would result in an average of one food delivery every 15 sec (since it would take an average of two presentations of the 5-sec search period to reach the 5-sec handling period for the short alternative), which is shorter than the 20-sec handling time for the long alternative.Suppose, however, that the search period is FI 7.5 sec rather than FI 5 sec. Now optimal diet theory predicts indifference between acceptance and rejection of the long alternative, because the average time to food is 20 sec whether the animal accepts the long alternative or returns to the search period. (In this case, if the long alternative were always rejected, it would take an average of two presentations of the 7.5-sec search period to reach the 5-sec handling period for the short alternative, which is a total
301Copyright 2008 Psychonomic Society, Inc.
Effects of reinforcer delay and variability in a successive-encounters procedure
JAMES E. MAZUR
Southern Connecticut State University, New Haven, ConnecticutPigeons responded in a successive-encounters procedure that consisted of a search period, a choice period, and a handling p...