Occasional Paper of the IUCN Species Survival Commission No. 55 About IUCN IUCN, International Union for Conservation of Nature, helps the world find pragmatic solutions to our most pressing environment and development challenges. IUCN's work focuses on valuing and conserving nature, ensuring effective and equitable governance of its use, and deploying nature-based solutions to global challenges in climate food and development. IUCN supports scientific research, manages field projects all over the world, and brings governments, NGOs, the UN and companies together to develop policy, laws and best practice. IUCN is the world's oldest and largest global environmental organization, with almost 1,300 government and NGO Members and more than 15,000 volunteer experts in 185 countries. IUCN's work is supported by almost 1,000 staff in 45 offices and hundreds of partners in public, NGO and private sectors around the world. http://www.iucn.org IUCN Species Survival Commission The Species Survival Commission (SSC) is the largest of IUCN's six volunteer commissions with a diverse global membership of nearly 10,000 experts. With biodiversity loss as one of the world's most pressing crises, and many species declining to critical levels, the SSC drives its efforts and expertise to conserving nature through positive and informed action. This is achieved through the strength of its dedicated sciencebased network of volunteer experts, who constitute the invaluable "power house" of the SSC. Web: http:// www.iucn.org/ssc IUCN Commission on Environment, Economics and Social Policy CEESP, the IUCN Commission on Environmental, Economic and Social Policy, is an interdisciplinary network of professionals whose mission is to act as a source of advice of the environmental, economic, social and cultural factors that affect natural resources and biological diversity, and to provide guidance and support towards effective policies and practices in environmental conservation and sustainable development. The designation of geographical entities in this book, and the presentation of the material, do not imply the expression of any opinion whatsoever on the part of IUCN, the European Union, the ACP Secretariat or the compilers concerning the legal status of any country, territory, or area, or of its authorities, or concerning the delimitation of its frontiers or boundaries. This publication has been produced with the support of the International Fur Federation. The views expressed in this publication do not necessarily reflect those of IUCN, the European Union, the ACP Secretariat, the International Fur Federation, or other participating organizations.