“…However, not all traits deemed to be ornaments have been found to play a vital role in female mate choice (Saether et al, 2000;Westneat, 2006;Takahashi et al, 2008;Shamble et al, 2009), causing researchers to re-evaluate the putative function of such secondary sexual traits. In lieu of an explanation requiring direct selection on male ornaments, increased support is surfacing for hypotheses suggesting that female mate choice is based upon combinations of traits or signal interactions (Kekäläinen et al, 2010;Grafe et al, 2012;Taylor et al, 2011) and/or upon motor performance, with ornamentation evolving secondarily in an enhancing role (see Byers et al, 2010;Barske et al, 2011;Hebets et al, 2011;Wilgers and Hebets, 2012). The former hypothesis proposes that separate signal components interact in various ways, dependent on signaling system, to alter female response.…”