In elementary particle theories the fine structure constant 2 / e c α = serves as the coupling constant for lepton interactions (QED), but is assumed to play no role in hadron interactions (QCD). However, experiments have long indicated an α spacing in the lifetimes of the long-lived threshold-state hadrons, and they also suggest an α-related mass structure. The relevance of α to hadron interactions is an experimental question, independent of theory. In the present paper we first make a detailed analysis of the experimental lifetime data. This analysis demonstrates that out of 156 particles with well-determined lifetimes τ, the 120 short excited-state lifetimes occur in α-spaced groups that cleanly sort out the s, c, b quark flavors. These 36 metastable lifetimes also exhibit a factor-of-three c-to-b "flavor structure" and a pervasive factor-of-two "hyperfine structure". We then trace out an α-defined set of mass quanta that tie together leptons and hadrons. Mass generation occurs via an initial "α-leap" from an electron pair to a "platform state" M, and then subsequent excitations by a dominant quantum X. The low-mass "MX octet" of particles-µ, p, τ, π, η, η', K, φ-is reproduced to an average accuracy of 0.4%, with no adjustable parameters except a small binding energy for hadronic pairs. Without the inclusion of lepton masses, the spectrum of hadron masses is difficult to understand. These reciprocal α-quantized results reinforce the reality of the spin 1/2 u, d, s, c, b quarks, and they also lead to the identification of a closely-related set of spinless mass quanta for the pseudoscalar mesons.