During the Devonian period, major continents/blocks were located in the southern hemisphere (e.g. Stampfli et al. 2002, 2013; Scotese 2014), with a major mountain range running on the northeastern margin of Laurussia to Gondwana: the Appalachian range (Fig. 1) (DeSantis 2010, Scotese 2014). This period of time is marked by one of the most significant modifications in palaeobiogeography with the precipitated end of the strong faunal endemism of the Emsian-Eifelian and the initiation of the Givetian-Frasnian cosmopolitanism (Oliver & Pedder 1979b). During the early Emsian to late Eifelian interval, three distinct faunal assemblages allow the definition of three different marine realms (Oliver & Pedder 1979a, 1979b, 1994; May 1995, 1997b), separated by various barriers. The Malvinokaffric Realm (MKR), located along the margins of Gondwana (Fig. 1), is relatively poor in corals and it is characterized by cold-water species (Oliver 1990;