“…Uplift rates based on paleobathymetries of marine sediments from across the Isthmus suggest that the majority of the volcanic arc was submerged up until very recently with bathyal sediments of latest Miocene age present in the Caribbean and Pacific of western Panama, the Caribbean of central Panama and eastern Panama (Coates et al, 1992;Collins, 1992Collins, , 1993Collins, , 1996Coates, 1999;Coates et al, 2000Coates et al, , 2003Coates et al, , 2004Coates et al, , 2005. Strong similarities between Pacific and Caribbean Mio-Pliocene biota (Chaisson and Ravelo, 2000;O'Dea et al, 2007b) and near-identical environmental conditions on either side of the Isthmus (Keigwin, 1978(Keigwin, , 1982Keller et al, 1989;Haug and Tiedemann, 1998;Haug et al, 2001) in the Early Pliocene imply the CAS was maintained in some form until the mid-Pliocene. Nevertheless, well-studied and extensive lacustrine sediments, shallow water reefs and mangroves, plus a continental-like mammal fauna in the Canal Zone of Panama demonstrate the presence of substantial subaerial land(s) in the middle Miocene (Whitmore and Stewart, 1965;Kirby and MacFadden, 2005;Johnson and Kirby, 2006;Retallack and Kirby, 2007;Kirby et al, 2008).…”