“…The presence or absence of alien trees in fynbos and encroaching native trees in grassland environments of South Africa have been linked to a wide range of nutrients, soil properties, and soil treatments, including pH, acid saturation, Mg, Ca, Mn, Cu, Zn, B, P, and N-fertilisation (Mills and Allen 2018;Mills et al 2017). Unpublished data collected from 25 diverse sites across South African Fynbos, Grassland and Savanna sites have shown that soils in sub-sites relatively poor in boron or relatively rich in phosphorus tend to be less wooded than adjacent sub-sites.…”