“…GCM prediction skill should be examined for various oceanic variables that are useful for biological forecasts, because skillful ocean predictions of quantities that drive biological models are needed to achieve skillful ecological forecasts. To date, oceanic prediction skill has been examined mainly for SST (e.g., Becker et al, 2014;Doi et al, 2019;Hervieux et al, 2019) including marine heatwaves (Jacox et al, 2022), sea-surface height (e.g., Widlansky et al, 2017;Long et al, 2021;Shin and Newman, 2021;Amaya et al, 2022), and upper-layer temperatures (e.g., Yeager et al, 2018;Doi et al, 2020), because these variables are important in describing physical climate variability and relatively easy to evaluate with observation-based products. However, for biological predictions, other variables (e.g., mixed-layer depth, upwelling, salinity, bottom temperature, vertical profiles of temperature and density) can also be important, as they impact nutrient availability and the habitat of marine species, and they may be associated with a higher degree of predictability (e.g., Siedlecki et al, 2016;Capotondi et al, 2019a).…”