In dryland ecosystems, the redistribution of rainfall runoff from bare soil to vegetated patches is fundamentally important to plant survival and the maintenance of hillslope functionality, sustaining vegetation in regions where annual rainfall alone would be insufficient (Dunkerley, 2002;Schlesinger et al., 1990). With soil infiltrability typically higher under vegetation cover than in bare soil areas (Assouline, 2004;Belnap, 2006), bare soil and vegetation patches create spatial mosaics of sources and sinks (see Figure 1a), where runoff generated in bare soil areas (sources) is trapped by downslope vegetated patches (sinks) (Dunkerley, 2002;Li et al., 2008;Moreno-de las Heras et al., 2012).More tortuous flowpaths that bypass vegetation can also form in drylands, for example, where plants grow on mounds (Michaelides et al., 2009;Parsons et al., 1996;Wainwright et al., 2000). The importance of vegetation resistance to flow in contributing to such tortuosity, however, is unclear, even though vegetated patches provide greater resistance to flow than bare areas (Dunne et al., 1991;Rossi et al., 2018). Higher infiltrability in vegetated patches relative to bare soil areas promotes "run-on" behavior (Thompson et al., 2011). Conversely, greater surface roughness-and thus greater resistance to flow-may divert runoff around vegetation, promoting "run-around" behavior and reducing the transport of resources to vegetated areas. On topographically smooth surfaces, the vegetation's effect on flowpaths must result from the interaction between its effects on the resistance and infiltration. Unraveling this interaction requires, at minimum, describing unsteady two-dimensional flows over spatially heterogeneous dryland hillslopes (Crompton et al., 2019).