Edible ectomycorrhizal mushrooms such as truffles (Tuber), chanterelles (Cantharellus cibarius Fr. and related species), porcinis (Boletus edulis Bull. and related species), saffron milk caps (Lactarius deliciosus [L.] Gray and related species), and matsutakes (Tricholoma matsutake [S. Ito & S. Imai] Singer and related species) have high economic value worldwide (Zambonelli & Bonito, 2012; Pérez-Moreno, Guerin-Laguette, Arzú, & Yu, 2020), with annual global sales estimated at several billion dollars. These mushrooms are harvested in forests or plantations for both domestic and export markets (Arora, 2008;Tsing, 2015, Pérez-Moreno et al., 2021. In Japan, imported matsutake mushrooms from countries such as China, USA, Canada, Morocco, Turkey, Mexico, and South Korea are valued at 80 to 100 million dollars annually (Aoki et al., 2022), which is equivalent to domestic T. matsutake production within Japan, mainly in Nagano and Iwate Prefectures (Yamanaka, . During the past 20 y, truffle imports to Japan have increased such that their economic value in Japan now equivalent to around 10% that of the matsutake mushroom import value (MAFF; https://www.maff.go.jp/e/index.html).To meet this increasing market demand, truffles have been harvested in tree plantations (Zambonelli, Iotti, & Murat, 2016). However, other edible mushrooms are largely harvested in forests because non-host and tree plantation cultivation techniques have not yet been established. Therefore, these mushrooms must be conserved in natural forests through controlled harvests or forest management to sustain moderate harvest levels for longer (decade-scale) periods (