“…The tidal flats served as a staging area and barrier between sea and land, but one in which essentially no fossil record is preserved. Understandably, Little (1990) begins his brief account of the development of terrestrial ecosystems with the reputed terrestrial forms in the Ordovician and does not consider earlier time (see also Gray and Shear, 1992). To our mind, Climactichnites is remarkable in showing a first attempt to cross this barrier.…”