Plant-plant interactions during seedling establishment can markate the rate of seedling emergence (Bergelson and Perry, edly affect the composition of pasture communities. This research examined the emergence, mortality, and early growth of four forage 1989). In some cases, shelter by neighboring plants was species commonly found in temperate northeastern U.S. pastures. necessary for seedling establishment (Ryser, 1993). Con-Species were selected based on functional group (grass vs. legume) versely, under less harsh environments earlier emerging and relative drought tolerance. Drought-tolerant species included individuals or individuals that emerge into a less com-'Penlate' orchardgrass (Dactylis glomerata L.) and 'Viking' birdsfoot petitive environment can have both greater biomass and trefoil (Lotus corniculatus L.), while drought-sensitive species ingreater probability of survival than later emerging seedcluded 'Basion' perennial ryegrass (Lolium perenne L.) and 'Will' lings (Ross and Harper, 1972; Miller, 1987; Bergelson white clover (Trifolium repens L.). Seeds were sown as monocultures, and Perry, 1989). This initial advantage appears to be as grass-legume binary mixtures, and as a complex, four-species mixan important feature of competition for light but not ture. Mixture complexity had only minor effects on seedling emerfor nutrients (Wilson, 1988). gence. However, legume mortality was significantly reduced in the complex compared with other mixtures in a year when high tempera-Increasing plant species diversity has the potential to ture and drought stress limited seedling establishment. In most cases increase both the productivity and stability of temperate there was a negative effect of neighbors on survival as evidenced by grasslands (Tilman, 1999; Hughes and Roughgarden, 2000; reduced clustering of surviving compared with emerged seedlings and Spehn et al., 2000). It, therefore, becomes essential to by a negative relationship between mortality rate and distance to the understand the dynamics of plant-plant interactions durnearest neighbor. However, in a drought year, perennial ryegrass ing multi-species pasture establishment. While much inmortality decreased as distance to the nearest neighbor decreased, formation is available on interplant interactions of masuggesting that survival was facilitated by the presence of neighbors. ture plants (Harper, 1977), relatively little information Although mixture complexity had significant effects on seedling emeris available on interactions during the seedling stage. gence and mortality, species composition in the binary and complex When seeds are sown in monoculture the fate of any mixtures could be predicted based on emergence and survival of monocultures. It appears that seedling emergence information particular individual is probably of minor importance gleaned from monocultures can be a useful tool for predicting initial as long as a vigorous stand is established. However, when species composition of more complex mixtures.