Please cite this article as: Smalley, I., O'Hara-Dhand, K., McLaren, S., Svircev, Z., Nugent, H., Loess and Bee-Eaters I: Ground properties affecting the nesting of European bee-eaters (Merops apiaster L.1758) in loess deposits, Quaternary International (2012), doi: 10.1016/j.quaint.2012.09.005. This is a PDF file of an unedited manuscript that has been accepted for publication. As a service to our customers we are providing this early version of the manuscript. The manuscript will undergo copyediting, typesetting, and review of the resulting proof before it is published in its final form. Please note that during the production process errors may be discovered which could affect the content, and all legal disclaimers that apply to the journal pertain.
AbstractThe European bee-eater (Merops apiaster L.1758) nests in tunnels in loess deposits. The properties of loess make it particularly suitable for tunnel nesting birds (a major factor is the metastable nature of the ground). The 'Heneberg Compromise' operates whereby the conflicting requirements of tunnel stability and ease of excavation dictate the optimum particle size for usable ground. The open structure of loess deposits, due to particle shape and airfall sedimentation, allows gas movement in nesting tunnels. It also allows local compaction during nest building which strengthens tunnel walls. The short range nature of the interparticle bonds in the ground material provides an almost ideal construction environment, ensuring a low plasticity index, which appears to be critical for tunnel building birds. Bee-eaters and sand martins dig tunnels in loess as 'primary nesters'.These loess tunnels are used by many 'secondary nesters'. The bee-eater is an efficient ecosystem engineer.